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The Previously unknown purpose of the "Ninth Chevron" is revealed, and ends up taking a team to an Ancient ship "Destiny", a ship built millions of years ago by the Ancients, used to seed Distant galaxies with Stargates. This team, led by Dr. Nicolas Rush and Colonel Everett Young, is trapped on the ship, unable to change its programmed mission. The team encounters new races, new technology and new enemies as the runaway ship travels to the far ends of the Universe.
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An attack on a secret off-world base by a rebel organization has stranded the remaining survivors on an Ancient ship "Destiny", a large unmanned ship launched millions of years ago. At the far end of the universe, Colonel Everett Young and Dr. Nicolas Rush now lead the new "Destiny" crew on a journey to discovering new worlds as well as encountering new alien creatures, in hope of finding a way back home. Morals are tested and tensions rise when Colonel Young's leadership is questioned all while Dr. Rush sacrifices the safety of the crew to pursue his own scientific ambitions. Along the way, they make new enemies and clash with old familiar ones, as they uncover the mystery behind Destiny's true purpose.